Publications (including submissions & preprints)
Miller, C.W., Kimball, R.T., Forthman, M. 2024. Evolution of multi-component weapons in the superfamily of leaf-footed bugs. Evolution, 78: 635–651. [PDF]
Forthman, M. 2024. First record of Pyrrhocoris apterus (Linnaeus 1758) (Hemiptera: Heteroptera: Pentatomidae) in California, U.S.A. Pan-Pac. Entomol., 100: 75–78. [Link]
Forthman, M., Phan, H., Miller, C.W., Kimball, R.T. (In press). Phylogenetic placement of the leaf-footed bug tribes Agriopocorini, Amorbini, and Manocoreini (Heteroptera: Coreidae) using ultraconserved elements. Zool. J. Linn. Soc., zlae024. [Link]
Forthman, M., Gordon, E.R.L., Kimball, R.T. 2023. Low hybridization temperatures improve target capture success of invertebrate loci. R. Soc. Open Sci., 10: 230307. [PDF]
Forthman, M., Downie, C., Miller, C.W., Kimball, R.T. 2023. Evolution of stridulatory mechanisms: vibroacoustic communication may be common in leaf-footed bugs and allies (Heteroptera: Coreoidea). R. Soc. Open Sci., 10: 221348. [PDF]
Adler, K., Schill, A.E.R., Stolberg, A.M., Miller, C.W., Forthman, M. 2022. First record of the bow-legged bug Hyalymenus subinermis (Heteroptera: Alydidae) in California, with description of the mimetic immature stages. Pan-Pac. Entomol., 98: 138–149. [Link]
Forthman, M., Miller, C.W., Kimball, R.T. 2022. Phylogenomic analysis with improved taxon sampling corroborates an Alydidae + Hydarinae + Pseudophloeinae clade (Heteroptera: Coreoidea: Alydidae, Coreidae). Org. Divers. Evol., 22: 669–679. [Link]
Forthman, M., Lara, R., Meeds, A.W., Rider, D.A. 2022. First record of Pellaea stictica (Dallas, 1851) (Hemiptera: Heteroptera: Pentatomidae) in California, U.S.A. Pan-Pac. Entomol., 98: 76–80. [Link]
Miller, C.D., Forthman, M., Miller, C.W., Kimball, R.T. 2022. Extracting “legacy loci” from an invertebrate sequence capture dataset. Zool. Scr., 51: 14–31. [Link]
Forthman, M., Braun, E.L., Kimball, R.T. 2022. Gene tree quality affects empirical coalescent branch length estimation. Zool. Scr., 51: 1–13. [Link]
Forthman, M., Gil-Santana, H.R. 2021. Two new species of Rhiginia Stål, 1859, with taxonomical notes on species in the “cruciata-group” of this genus and an updated key to the New World genera of Ectrichodiinae (Heteroptera, Reduviidae). Zootaxa, 4952: 201–234. [Link]
Forthman, M. 2021. Two new species of Abelocephala (Heteroptera: Reduviidae) from Taiwan. Zootaxa, 4920: 278–286. [Link]
Forthman, M., Miller, C.W., Kimball, R.T. 2020. Phylogenomics of the leaf-footed bug subfamily Coreinae (Hemiptera: Coreidae). Insect Syst. Divers., 4: 2. [Link]
Emberts, Z., St. Mary, C.M., Howard, C.C., Forthman, M., Bateman, P.W., Somjee, U., Hwang, W.S., Li, D., Kimball, R.T., Miller, C.W. 2020. The evolution of autotomy in leaf-footed bugs. Evolution, 74: 897–910. [PDF]
Matzinger, E.E., Forthman, M. 2019. Identification key to the genera of the tribe Gonocerini (Insecta: Hemiptera: Coreidae). UF Journal of Undergraduate Research, 21: 1–7. [PDF]
Forthman, M., Miller, C.W., Kimball, R.T. 2019. Phylogenomic analysis suggests Coreidae and Alydidae (Hemiptera: Heteroptera) are not monophyletic. Zool. Scr., 48: 520–534. [PDF]
Kieran, T.J., Gordon, E., Forthman, M., Hoey-Chamberlain, R., Kimball, R.T., Faircloth, B.C., Weirauch, C., Glenn, T.C. 2019. Insights from an ultraconserved element bait set designed for hemipteran phylogenetics integrated with genomic resources. Mol. Phylogenet. Evol., 130: 297–303. [PDF]
Chen, D., Braun, E.L., Forthman, M., Kimball, R.T., Zhang, Z. 2018. A simple strategy for recovering ultraconserved elements, exons, and introns from low coverage shotgun sequencing of museum specimens: placement of the partridge genus Tropicoperdix within the Galliformes. Mol. Phylogenet. Evol., 129: 304–314. [PDF]
Forthman, M., Weirauch, C. 2018. Phylogenetic comparative analysis supports aposematic colouration–body size association in millipede assassins (Hemiptera: Reduviidae: Ectrichodiinae). J. Evol. Biol., 31: 1071–1078. [PDF]
Forthman, M., Weirauch, C. 2017. Millipede assassins and allies (Heteroptera: Reduviidae: Ectrichodiinae, Tribelocephalinae): total evidence phylogeny, revised classification and evolution of sexual dimorphism. Syst. Entomol. 42: 575–595. [PDF]
Weirauch, C., Forthman, M., Grebenikov, V. Baňař, P. 2017. From Eastern Arc Mountains to extreme sexual dimorphism: systematics of the enigmatic assassin bug genus Xenocaucus (Hemiptera: Reduviidae: Tribelocephalinae). Org. Divers. Evol. 17: 421–445. [PDF]
Forthman, M., Weirauch, C. 2016. Phylogenetics and biogeography of the endemic Madagascan millipede assassin bugs (Heteroptera: Reduviidae: Ectrichodiinae). Mol. Phylogenet. Evol. 100: 219–233. [PDF]
Forthman, M., Chłond, D., Weirauch, C. 2016. Taxonomic monograph of the endemic millipede assassin bug fauna of Madagascar (Heteroptera: Reduviidae: Ectrichodiinae). B. Am. Mus. Nat. Hist., 400: 1–152. [Link]
Zhang, J., Gordon, E., Forthman, M., Hwang, W.S., Walden, K., Swanson, D., Johnson, K.P., Meier, R., Weirauch, C. 2016. Evolution of the assassin’s arms: insights from a phylogeny of combined transcriptomic and ribosomal DNA data (Heteroptera: Reduvioidea). Sci. Rep. 6: 22177. [PDF]
Weirauch, C., Berenger, J.M., Berniker, L., Forero, D., Forthman, M., Frankenberg, S., Freedman, A., Gordon, E., Hoey-Chamberlain, R., Hwang, W.S., Michael, A., Udah, O., Watson, C., Zhang, G., Zhang, J. 2014. An illustrated identification key to assassin bug subfamilies and tribes (except Emesinae). Can. J. Arthropod Ident., No. 26. [Link]
Forthman, M., Weirauch, C. 2012. Toxic associations: a review of the predatory behaviors of millipede assassin bugs (Hemiptera: Reduviidae: Ectrichodiinae). Eur. J. Entomol., 109: 147–153. [PDF]
Forthman, M. 2024. First record of Pyrrhocoris apterus (Linnaeus 1758) (Hemiptera: Heteroptera: Pentatomidae) in California, U.S.A. Pan-Pac. Entomol., 100: 75–78. [Link]
Forthman, M., Phan, H., Miller, C.W., Kimball, R.T. (In press). Phylogenetic placement of the leaf-footed bug tribes Agriopocorini, Amorbini, and Manocoreini (Heteroptera: Coreidae) using ultraconserved elements. Zool. J. Linn. Soc., zlae024. [Link]
Forthman, M., Gordon, E.R.L., Kimball, R.T. 2023. Low hybridization temperatures improve target capture success of invertebrate loci. R. Soc. Open Sci., 10: 230307. [PDF]
Forthman, M., Downie, C., Miller, C.W., Kimball, R.T. 2023. Evolution of stridulatory mechanisms: vibroacoustic communication may be common in leaf-footed bugs and allies (Heteroptera: Coreoidea). R. Soc. Open Sci., 10: 221348. [PDF]
Adler, K., Schill, A.E.R., Stolberg, A.M., Miller, C.W., Forthman, M. 2022. First record of the bow-legged bug Hyalymenus subinermis (Heteroptera: Alydidae) in California, with description of the mimetic immature stages. Pan-Pac. Entomol., 98: 138–149. [Link]
Forthman, M., Miller, C.W., Kimball, R.T. 2022. Phylogenomic analysis with improved taxon sampling corroborates an Alydidae + Hydarinae + Pseudophloeinae clade (Heteroptera: Coreoidea: Alydidae, Coreidae). Org. Divers. Evol., 22: 669–679. [Link]
Forthman, M., Lara, R., Meeds, A.W., Rider, D.A. 2022. First record of Pellaea stictica (Dallas, 1851) (Hemiptera: Heteroptera: Pentatomidae) in California, U.S.A. Pan-Pac. Entomol., 98: 76–80. [Link]
Miller, C.D., Forthman, M., Miller, C.W., Kimball, R.T. 2022. Extracting “legacy loci” from an invertebrate sequence capture dataset. Zool. Scr., 51: 14–31. [Link]
Forthman, M., Braun, E.L., Kimball, R.T. 2022. Gene tree quality affects empirical coalescent branch length estimation. Zool. Scr., 51: 1–13. [Link]
Forthman, M., Gil-Santana, H.R. 2021. Two new species of Rhiginia Stål, 1859, with taxonomical notes on species in the “cruciata-group” of this genus and an updated key to the New World genera of Ectrichodiinae (Heteroptera, Reduviidae). Zootaxa, 4952: 201–234. [Link]
Forthman, M. 2021. Two new species of Abelocephala (Heteroptera: Reduviidae) from Taiwan. Zootaxa, 4920: 278–286. [Link]
Forthman, M., Miller, C.W., Kimball, R.T. 2020. Phylogenomics of the leaf-footed bug subfamily Coreinae (Hemiptera: Coreidae). Insect Syst. Divers., 4: 2. [Link]
Emberts, Z., St. Mary, C.M., Howard, C.C., Forthman, M., Bateman, P.W., Somjee, U., Hwang, W.S., Li, D., Kimball, R.T., Miller, C.W. 2020. The evolution of autotomy in leaf-footed bugs. Evolution, 74: 897–910. [PDF]
Matzinger, E.E., Forthman, M. 2019. Identification key to the genera of the tribe Gonocerini (Insecta: Hemiptera: Coreidae). UF Journal of Undergraduate Research, 21: 1–7. [PDF]
Forthman, M., Miller, C.W., Kimball, R.T. 2019. Phylogenomic analysis suggests Coreidae and Alydidae (Hemiptera: Heteroptera) are not monophyletic. Zool. Scr., 48: 520–534. [PDF]
Kieran, T.J., Gordon, E., Forthman, M., Hoey-Chamberlain, R., Kimball, R.T., Faircloth, B.C., Weirauch, C., Glenn, T.C. 2019. Insights from an ultraconserved element bait set designed for hemipteran phylogenetics integrated with genomic resources. Mol. Phylogenet. Evol., 130: 297–303. [PDF]
Chen, D., Braun, E.L., Forthman, M., Kimball, R.T., Zhang, Z. 2018. A simple strategy for recovering ultraconserved elements, exons, and introns from low coverage shotgun sequencing of museum specimens: placement of the partridge genus Tropicoperdix within the Galliformes. Mol. Phylogenet. Evol., 129: 304–314. [PDF]
Forthman, M., Weirauch, C. 2018. Phylogenetic comparative analysis supports aposematic colouration–body size association in millipede assassins (Hemiptera: Reduviidae: Ectrichodiinae). J. Evol. Biol., 31: 1071–1078. [PDF]
Forthman, M., Weirauch, C. 2017. Millipede assassins and allies (Heteroptera: Reduviidae: Ectrichodiinae, Tribelocephalinae): total evidence phylogeny, revised classification and evolution of sexual dimorphism. Syst. Entomol. 42: 575–595. [PDF]
Weirauch, C., Forthman, M., Grebenikov, V. Baňař, P. 2017. From Eastern Arc Mountains to extreme sexual dimorphism: systematics of the enigmatic assassin bug genus Xenocaucus (Hemiptera: Reduviidae: Tribelocephalinae). Org. Divers. Evol. 17: 421–445. [PDF]
Forthman, M., Weirauch, C. 2016. Phylogenetics and biogeography of the endemic Madagascan millipede assassin bugs (Heteroptera: Reduviidae: Ectrichodiinae). Mol. Phylogenet. Evol. 100: 219–233. [PDF]
Forthman, M., Chłond, D., Weirauch, C. 2016. Taxonomic monograph of the endemic millipede assassin bug fauna of Madagascar (Heteroptera: Reduviidae: Ectrichodiinae). B. Am. Mus. Nat. Hist., 400: 1–152. [Link]
Zhang, J., Gordon, E., Forthman, M., Hwang, W.S., Walden, K., Swanson, D., Johnson, K.P., Meier, R., Weirauch, C. 2016. Evolution of the assassin’s arms: insights from a phylogeny of combined transcriptomic and ribosomal DNA data (Heteroptera: Reduvioidea). Sci. Rep. 6: 22177. [PDF]
Weirauch, C., Berenger, J.M., Berniker, L., Forero, D., Forthman, M., Frankenberg, S., Freedman, A., Gordon, E., Hoey-Chamberlain, R., Hwang, W.S., Michael, A., Udah, O., Watson, C., Zhang, G., Zhang, J. 2014. An illustrated identification key to assassin bug subfamilies and tribes (except Emesinae). Can. J. Arthropod Ident., No. 26. [Link]
Forthman, M., Weirauch, C. 2012. Toxic associations: a review of the predatory behaviors of millipede assassin bugs (Hemiptera: Reduviidae: Ectrichodiinae). Eur. J. Entomol., 109: 147–153. [PDF]